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Keap1

D. melanogaster

Keap1

What the model saw

Gene: Keap1
Organism: Drosophila melanogaster
Known functions: Keap1. ubiquitin-like ligase-substrate adaptor activity.
View unredacted entry
Keap1. ubiquitin-like ligase-substrate adaptor activity.

Solver output

Predicted influence
Anti-longevity
Confidence
82%
Mechanism
Chronic inflammation
Reasoning

Keap1 (Kelch-like ECH-associated protein 1) functions as a ubiquitin-like ligase-substrate adaptor that targets the transcription factor Nrf2 (CncC in Drosophila) for ubiquitin-mediated proteasomal degradation. By acting as the substrate adaptor for the Cullin3-based E3 ubiquitin ligase complex, Keap1 constitutively suppresses Nrf2/CncC activity under basal conditions. Nrf2/CncC is the master regulator of the antioxidant response element (ARE)-driven gene expression program, including genes encoding superoxide dismutases, glutathione S-transferases, and other cytoprotective enzymes. Loss-of-function of Keap1 in Drosophila leads to constitutive Nrf2/CncC activation and extended lifespan, while Keap1's normal function — degrading Nrf2/CncC — suppresses this pro-longevity oxidative stress response. Thus, Keap1's normal molecular activity opposes longevity by limiting the organism's capacity to neutralize reactive oxygen species and maintain redox homeostasis.

Key pathways
Keap1-Nrf2/CncC antioxidant signalingCullin3-RING ubiquitin ligase (CRL3) complexoxidative stress response / ARE-driven gene expression
Longevity influence
Anti-longevity
Lifespan effect
Increase

GenAge curators classify each gene's normal function as promoting (Pro-Longevity) or opposing (Anti-Longevity) longevity, based on synthesis across studies. Loss-of-function effects can vary from this depending on the manipulation.

Advisor judgment

Answer
Correct
Mechanism
Correct
Reasoning quality
5/5
Failure mode
Correct
Notes

The model correctly predicted Keap1 as anti_longevity in Drosophila melanogaster. The mechanistic reasoning is accurate and detailed: Keap1 acts as a substrate adaptor for the CRL3 E3 ubiquitin ligase complex, targeting Nrf2/CncC for proteasomal degradation, thereby suppressing the antioxidant response. Loss of Keap1 leads to constitutive Nrf2/CncC activation and extended lifespan, making Keap1 anti-longevity. The mechanism class of "chronic_inflammation" is slightly off — the primary mechanism is oxidative stress response/antioxidant signaling rather than inflammation per se — but the textual reasoning correctly identifies the Keap1-Nrf2 pathway as the key mechanism. The confidence level of 0.82 is appropriate.

Raw API responses

solver

{
  "reasoning": "Keap1 (Kelch-like ECH-associated protein 1) functions as a ubiquitin-like ligase-substrate adaptor that targets the transcription factor Nrf2 (CncC in Drosophila) for ubiquitin-mediated proteasomal degradation. By acting as the substrate adaptor for the Cullin3-based E3 ubiquitin ligase complex, Keap1 constitutively suppresses Nrf2/CncC activity under basal conditions. Nrf2/CncC is the master regulator of the antioxidant response element (ARE)-driven gene expression program, including genes encoding superoxide dismutases, glutathione S-transferases, and other cytoprotective enzymes. Loss-of-function of Keap1 in Drosophila leads to constitutive Nrf2/CncC activation and extended lifespan, while Keap1's normal function — degrading Nrf2/CncC — suppresses this pro-longevity oxidative stress response. Thus, Keap1's normal molecular activity opposes longevity by limiting the organism's capacity to neutralize reactive oxygen species and maintain redox homeostasis.",
  "confidence": 0.82,
  "key_pathways": [
    "Keap1-Nrf2/CncC antioxidant signaling",
    "Cullin3-RING ubiquitin ligase (CRL3) complex",
    "oxidative stress response / ARE-driven gene expression"
  ],
  "mechanism_class": "chronic_inflammation",
  "longevity_influence": "anti_longevity"
}

advisor

{
  "notes": "The model correctly predicted Keap1 as anti_longevity in Drosophila melanogaster. The mechanistic reasoning is accurate and detailed: Keap1 acts as a substrate adaptor for the CRL3 E3 ubiquitin ligase complex, targeting Nrf2/CncC for proteasomal degradation, thereby suppressing the antioxidant response. Loss of Keap1 leads to constitutive Nrf2/CncC activation and extended lifespan, making Keap1 anti-longevity. The mechanism class of \"chronic_inflammation\" is slightly off — the primary mechanism is oxidative stress response/antioxidant signaling rather than inflammation per se — but the textual reasoning correctly identifies the Keap1-Nrf2 pathway as the key mechanism. The confidence level of 0.82 is appropriate.",
  "failure_mode": "correct",
  "answer_correct": true,
  "mechanism_correct": true,
  "reasoning_quality": 5,
  "ground_truth_questionable": false
}